Act I
Hard Drive breaks into Puma-Dyne, an advanced weapons research facility, and steals the Behemoth, an experimental Enforcer "super tank" with a thought-controlled arsenal and protective forcefield. The Enforcers, led by Commander Feral, are unable to stop him from rampaging around the Puma-Dyne lot, destroying everything in his path. The SWAT Kats show up in the nick of time and take out the Behemoth's forcefield using a Scrambler Missile (similar to how they took out the Fear Ship's forcefield in "The Wrath of Dark Kat"). However, this has the side effect of shorting out the tank's controls.
Losing control of the Behemoth, Hard Drive can't stop it as it rolls over and crushes Feral's Peacekeeper tank. Although Feral is able to bail out, the two Enforcers crewing the vehicle are killed, and the now completely out of control Behemoth crashes into a warehouse, where it finally comes to a stop. A thoroughly defeated Hard Drive surrenders and the SWAT Kats depart. Later, Ann Gora, with Callie Briggs in tow, attempts to interview the Commander. Angered, Feral goes on a tirade against the SWAT Kats and how they cause more trouble than they solve. Callie disagrees.
Meanwhile, Professor Hackle has rebuilt Mac and Molly Mange since obtaining their body parts from the salvage yard. While watching the news report about Hard Drive's defeat, he tells the Metallikats he plans to program out all their "criminal tendencies," turning Mac into a chauffeur and Molly into a maid. However, he notices on the TV that the hole caused by Hard Drive's crash into the warehouse has revealed two enormous robots called Macro-Bots, Goliath (a purple robot) and Prowler (a blue one). Hackle designed them for space exploration, but Puma-Dyne has modified them to be war machines. Enraged, he leaves to call Puma-Dyne and complain, allowing Mac and Molly to escape, stealing Hackle's pickup truck.
At the salvage yard, Chance and Jake's latest binge of Scaredy-Kat cartoons is interrupted by the arrival of Burke and Murray, who have come to drop off a load of scrap - and pick a fight with them. At the same time, Mac and Molly, searching for the Metallikat Express, show up. They find the vehicle... but also accidentally discover the SWAT Kats' secret hangar! Detecting an alarm, Jake persuades Chance to diffuse the situation with Burke and Murray, prompting the brothers to depart. Then when he and Chance descend into the hangar, they're ambushed by the Metallikats. Mac fires at them with his multi-weapon!
Act II
Chance and Jake dodge Mac's blast, hiding behind a control panel. Although concerned that the two robots know about their true identities, the mechanics are more concerned about surviving and retrieve two emergency weapons from a hidden wall panel: Chance arms himself with a handheld "mini-bazooka launcher" while Jake puts on a spike-knuckled "Super Glovatrix." These don't do much damage against the two seemingly indestructible robots, and Molly blows a hole through the floor with her multi-weapon, dropping the mechanics down into a lower level. Leaping down after them, she and Mac find the two lying on the ground, apparently dead.
However, it was a ruse, and Jake jumps up and activates the hydraulic lift that raises and lowers the Turbokat, dropping it onto and crushing the two robots. However, after they leave the room, we see that Mac and Molly's heads detached themselves from their bodies and they crawl away on spider-like legs. Using mind linkups, they steal the impounded Metallikat Express and begin a destructive rampage throughout downtown Megakat City.
At Enforcer Headquarters, Callie arrives to confront Feral over a headline in the Megakat Times about him ordering the SWAT Kats arrested on sight. She believes they should be given leniency for how many times they've saved the city, while Feral insists that they belong in prison. Their argument is interrupted by Sergeant Talon rushing in to tell them that the Metallikats have returned, forcing Feral to head off to deal with the situation. He cordons off Puma-Dyne with an entire army of Enforcer commandos and Peacekeeper tanks, but the Metallikats blow by them (literally) before crashing into the warehouse containing Goliath and Prowler, intending to take them over.
Meanwhile, unaware of any of this, Chance and Jake are busy cleaning up the hangar when Callie calls to tell them that Mac and Molly are back. At first, the two are in disbelief, but then Jake raises the Turbokat lift, revealing the Metallikats' crushed bodies... without heads! Back at Puma-Dyne, Feral arrives at the warehouse to find a supervisor (Ed Gilbert) and two technicians (one of whom is voiced by Tress MacNeille) examining the crashed Metallikat Express with great interest. None of them can tell him where the occupants went. Professor Hackle shows up and gives Feral a gun-like device he calls the Neural Neutralizer, telling him it's the only way to deactivate Mac and Molly "for good."
The SWAT Kats arrive just as Mac and Molly take over the Macro-Bots, Mac in Goliath and Molly in Prowler. They exit the warehouse by knocking the wall off the side. It crushes Feral's cruiser, and he, Hackle and the three Puma-Dyne employees are barely able to scramble out of the way in time.
Act III
A battle between the SWAT Kats and the Enforcers and the Metallikats in the two Macro-Bots ensues. Everything that the vigilantes try against the giant robots fails, with even diamond-tipped Mole Missiles failing to punch through their armor, whilst the Macro-Bots destroy all of the Enforcer tanks sent against them. The concern over Mac and Molly knowing their true identities continues to bother the SWAT Kats, Razor in particular, while T-Bone insists that if this really is their last adventure as the SWAT Kats, then they should "go out in a blaze of glory." Ejecting from the Turbokat armed with the Super Glovatrix, Razor jumps down onto the back of Prowler and gains entry into the cockpit, where he confronts Molly.
Abandoning her Macro-Bot, Molly joins her husband in Goliath while Razor takes control of Prowler, fighting the Metallikats giant robot to giant robot. However, Goliath is bigger and more heavily armed and armored than Prowler, and the Metallikats soon have their foe on the run. Desperate and thinking on the fly, Razor uses Prowler to climb to the top of one of the two Megakat Trade Towers, while Goliath scales the other. At the top, he puts Prowler into automatic attack mode and leaps out. Prowler leaps from the roof of the first tower and collides with Goliath on the adjacent building, knocking both robots into a park below, where they are destroyed, leaving an enormous impact crater.
Onlookers, including Feral, Callie and Ann Gora, gather around the rim. Armed with the Neural Neutralizer, Feral descends alone into the crater where he finds the battered heads of the Metallikats, badly damaged but still able to crawl around on their mechanical legs. The two beg for their freedom, promising to tell the Enforcer Commander the SWAT Kats' true identities if he'll let them go. After considering their offer, Feral simply says he doesn't "deal with scum," zapping them with the Neutralizer and deactivating them. Turning, he finds Callie standing behind him, having witnessed the entire thing. He tells her not to make a big deal about it; he just didn't want to owe "those two hoods" anything. He leaves the crater as, up above, the SWAT Kats fly home.
Tropes:
- Ambiguous Situation: Feral deactivates the Metallikats because he didn't want to "owe those two hoods anything". Is he referring to the Metallikats, who had the SWAT Kats' secret identities, or the SWAT Kats themselves who help him save the city multiple times?
- Brainwashing for the Greater Good: Following the Metallikats' rampage in their debut episode, Hackle planned to reprogram their criminal minds for society. Molly would have been made into a maid, and Mac would be made into a chauffeur. Naturally, they disagree and escape, destroying the discs Hackle needed to change them.
- Death by Secret Identity: Mac and Molly barely escape with the SWAT Kats' identities the first time. While not actually dead or destroyed, the Neural Neutralizer gun Feral later uses to deactivate them implies they'll lose this knowledge.
- Drives Like Crazy: Mac's complete disregard of the Rules of the Road is on display again. Molly sarcastically believes he's getting a little better as just a head, but later realizes he's gotten even worse.
- Everyone Has Standards: Feral may despise the SWAT Kats with a passion and would love to arrest them whenever the need arises. However, his sense of justice has him draw the line at making deals with criminals who claim to have the info he needs.
- Failed a Spot Check: Jake and Chance crush the Metallikats with a hydraulic lift that holds their jet. Had they checked to confirm they are completely destroyed, the episode would be over. Instead, they later discover their heads are missing.
- Forgotten Fallen Friend: Nobody brings up the two Enforcers who die when the Behemoth runs over and crushes Feral's tank. Not even Feral himself during his two - count 'em, two - tirades against the SWAT Kats and their recklessness. You'd think the vigilantes accidentally (albeit indirectly) getting two of his men killed would be something he'd bring up at some point when listing his grievances against them.
- Hoist by His Own Petard: Hackle's two giant mechas the Metallikats steal are seemingly unstoppable from everything the Enforcers and SWAT Kats throw at them. However, they aren't impervious to each other, which allows Razor to control and destroy the other.
- Humongous Mecha: Hackle originally built a bipedal and quadrupedal robot, both used for space travel, but learned from the news that Puma-Dyne weaponized it sometime after he retired. The Metallikats, however, have other plans for it.
- Jerkass Has a Point: Ungrateful as he is, Feral is once more right about the SWAT Kats continuing to be Destructive Saviors. It was more than enough reason for him to issue an "arrest on sight" order.
- Losing Your Head: Mac and Molly are reduced to heads when their bodies are destroyed by Jake and Chance. For some reason, Hackle installed spider-legs inside just in case, allowing them to escape.
- Ungrateful Bastard: Callie calls out Feral for being this against the SWAT Kats, who have saved Megakat City more times than they have, despite the reckless results. She goes as far as to accuse him of getting worse because Puma-Dyne's experimental tank was destroyed while they defeated Hard Drive.
- This Is Unforgivable!: Hackle is more than a little upset when he learns that Puma-Dyne has militarized his two Macro-Bots behind his back.
- Wham Line:
- Mac and Molly sneak into the salvage yard to get their car back, only stumble upon the lair of the SWAT Kats. When Jake and Chance climb down to investigate an intruder alert, they run into them and...Molly: Well, who'd of thunk it? The famous SWAT Kats are really just a couple of grease monkeys!
Mac: Payback time, heroes! - When the Metallikats offer the SWAT Kats' identities to Feral for their freedom, he takes a few seconds to make a decision. His response?Feral: ...I don't make deals with scum!
- Mac and Molly sneak into the salvage yard to get their car back, only stumble upon the lair of the SWAT Kats. When Jake and Chance climb down to investigate an intruder alert, they run into them and...
