It is shown that Nars is rather abusive to Fleet when it comes to competition. And Fleet's fellow teammate Magril always whispers to Fleet because she's utterly terrified of Nars. It almost makes you wonder just how Nars treated the rest of his team.
Did the Head Wolf ever try to kill other pups like Daria? Or possibly succeed in doing so?
Kate and Humphrey seem fairly familiar with human inventions, and can read English signs despite their packs being relatively secluded from humans. Given that the setting is a national park, though, it could explain some familiarity with humans.