The monstrous Mabus, ruler of the Gua, has guided them on conquests that inevitably become genocides, exterminating all life on multiple planets. When he arrives on Earth, Mabus kills any ranking officer who dares to defy his orders while attempting to push for the Second Wave to kill millions of human civilians. Mabus, when his husk is destroyed, begins bodyjacking other humans, painfully obliterating their minds when he burns them out, eventually taking over the heroine Jordan and hunting down and wiping out entire resistance cells. He then even rapes the hero Cade Foster in Jordan's body to conceive a child that will be his perfect host. Mabus attempts to bring about an invasion, intending on killing every human being alive, obsessed with his own power and glory and caring for nothing else—not even the other Gua.
Cain is Mabus's second-in-command and second only to his leader in depravity among the Gua. Serving as the sympathetic Gua Joshua's replacement, Cain kidnaps Cade to subject him to horrible mental torture to plumb the Book of Nostradamus from his mind, apathetic to his survival through the process. Cain murders several people along the way, from using a cop as a human bomb to murdering one of his own minions to pay for another's failure, and later tortures a man to death in order to find the location of resistance base Raven Nation. Cain's attempt to slaughter Raven Nation fails, but he gets away with murdering one of their leaders in the process out of spite. Cain's ultimate goal is to launch the Second Wave, which would kill 19 million initially and millions later, remaining suavely sadistic the whole while he's onscreen.
"Blind Witness": Dr. Reid is a twisted Gua scientist who believes humans are nothing but livestock. Running a hospital, Reid takes the human patients and performs agonizing experiments on them that either kill them or result in Reid having them disposed of when he's finished. Reid grafts new parts to them, or carves them up to find out what makes them tick, before attempting to murder his only success, a former blind woman, along with Cade himself.
"Unearthed": Logan Bishop poses as one of the heads of an archaeological dig to find the legendary Hammer of Thor, in actuality a Gua artifact. Working directly on orders from Mabus, Bishop murders the leader of the dig once his use wears out by snapping his neck and tries to torture the location of other agents out of Eddie. Bishop's true goal is to use the Hammer of Thor to annihilate all life on Earth, even sadistically mocking the heroes that they'll be at the forefront of the slaughter he'll unleash.
