Original 8
Filia was once an ordinary schoolgirl, before her family was massacred by the Medici mafia. Filia herself only survived due to the intervention of the Parasite Samson, who bonded to her and replaced her hair. This left Filia with amnesia, and she seeks out the Skull Heart so that she can wish to restore her memories.
After Samson fends off a lecherous Medici goon, the two fight Parasoul and Cerebella as part of Samson's strategy to gain a lead on the Skull Heart by looking for trouble. This routine is interrupted by a blast from Filia's past; her old friend Carol, now transformed into an anti-Skullgirl weapon called Painwheel.
After a fight with Ms. Fortune, Filia runs into Painwheel again in the Grand Cathedral, where she's confronting one of her abductors, Valentine. Filia then fights Valentine and Double as a Dual Boss, and Double tells her that the Skull Heart can indeed restore her memories. Filia descends into the cathedral's catacombs to face off against the Skullgirl herself, Marie. Marie reveals that she has met Filia before... and tried to kill her because Filia herself is a Medici, though her family wanted nothing to do with the mafia business.
After Filia defeats Marie, the Skull Heart is hers to wish on... but instead of wishing for her memories, she wishes for Painwheel to be able to live a normal life. This is granted, but because the wish isn't quite pure, Filia will eventually become the next Skullgirl. The Skull Heart tells her to make the most of the time she has left, and she and Carol return to school... for the moment.
Cerebella is a circus performer/mafia goon rescued as a baby by Vitale Medici, who she sees as a father and utterly adores. After one show, Vitale has a new job for her: track down Nadia Fortune and retrieve the Life Gem she stole at any cost. Cerebella happily agrees.
She meets and fights Filia before finding Ms. Fortune in Little Innsmouth. Cerebella demands the return of the Life Gem, but Ms. Fortune says it won't be that easy, and they fight. Afterwards, Ms. Fortune reveals that she literally can't return the Life Gem even if she wanted to, because it fused with her body. Cerebella is stymied by this, but resolves to capture Ms. Fortune and bring her back to Vitale so he can figure something out.
Once there, she has to fight Valentine and Double (taking the form of Cerebella) in order to get to Ms. Fortune, because Valentine wants to experiment with Ms. Fortune's immortality. After shooing off the competition, Double tells her that Ms. Fortune has entered the catacombs and intends to claim the Skull Heart, encouraging Cerebella to wish on it instead.
Down in the catacombs, Cerebella finds Marie instead of Ms. Fortune. And while Marie acknowledges Cerebella's pure heart, she's still a Medici and so Marie wants her dead. After defeating Marie, Ms. Fortune comes in to snatch the Skull Heart (Cerebella ignores it), but is caught by Cerebella's hat Living Weapon, Vice Versa. Ms. Fortune declares her intent to use her wish to revive her friends and take revenge on the Medici mafia... perhaps not the wisest thing to say to Cerebella, who considers them her family. Cerebella uses Vice Versa's strength to crush Ms. Fortune until she becomes a new Life Gem.
Vitale is happy with this outcome... but Cerebella is troubled about having to kill someone so horribly.
Peacock is a young girl rebuilt by by Anti-Skullgirl Lab 8 with cybernetic enhancements that essentially make her a living cartoon. When the Skullgirl is announced to be active, she goes out to face her, fighting Filia, Ms. Fortune, and Parasoul in the process- much to her annoyance, since she'd really like to get the job done in time to catch Annie, Girl of the Stars.
Unfortunately, this is not to be. The Skullgirl attacks Lab 8 after Valentine opened the way for her. Dr. Avian, the brains behind Peacock's weapons, is severely injured (and dies after talking to Peacock) and the rest are scattered. Peacock recognizes the Skullgirl as Marie, but before Peacock can chase after her, Double interrupts and turns into Peacock for a Mirror Match. Peacock finally gets to the cathedral to fight Valentine and Double together for what they did to Lab 8.
With all the distractions out of the way, Peacock finally reunites with Marie. The two were once childhood friends, and Marie became the Skullgirl to take revenge on the slavers who mutilated Patricia before she became Peacock. Unfortunately, Peacock can't just let Marie go even if she does agree with the cause of murdering slavers. The Skullgirl is a monster that kills everyone in her path, and Peacock can't guarantee that Marie wouldn't wig out and start killing innocents after the Medici were finished.
After the battle, Peacock destroys the Skull Heart and has one last heartfelt conversation with Marie, reassuring her that she still considers Marie a friend, and promising to take up the cause of Medici genocide in her stead. The story ends with her about to face off against the Medicis' top enforcer, Black Dahlia.
Parasoul is the princess of the Canopy Kingdom, commander of the Black Egrets, the most effective law enforcement officer in the entire country, and older sister to Umbrella. Parasoul, unlike most others, has no desire to use the Skull Heart as the last Skullgirl was her mother Queen Nancy- and Nancy almost destroyed the world. Naturally, Parasoul doesn't want to repeat the same mistake.
When the latest Skullgirl appears, Parasoul is taking Umbrella to the prison to see the various criminals Parasoul had captured, but this and their subsequent trip to get ice cream is cut short when Parasoul hears the news on the Skullgirl. Umbrella wants to help, but Parasoul doesn't want the young girl to get involved... and Umbrella wasn't an implemented fighter. Umbrella gets pissed by Parasoul constantly putting The Needs of the Many ahead of her, but she doesn't change Parasoul's mind. She then appears to see her mom and runs off... but remember what I said earlier about her mom having turned into the Skullgirl and died?
Meanwhile, back with Parasoul, she fights Ms. Fortune, Filia, and Painwheel. She's then blocked from approaching the Skullgirl by Umbrella, who believes that the Skull Heart is her mom. Valentine takes Umbrella away to experiment with her unique connection to the Skull Heart, while Double fights Parasoul as a Mirror Match. Double then confirms to Parasoul that Umbrella, uniquely enough, was born while her mother was in the process of turning into the Skullgirl. This gave her an innate connection to its power... and unfortunately, an innate weakness to its temptation. And if she were to become a Skullgirl, she'd be a Skullgirl even more powerful than Nancy... and did I mention that Nancy almost destroyed the world?
Parasoul rushes off to the Grand Cathedral, fights off Double and Valentine, and confronts Marie. Marie didn't do anything to Umbrella (who reminded her of Patricia), but she didn't need to; just being near the Skull Heart drained Umbrella's life force. Parasoul fights Marie but cannot destroy the Skull Heart as Umbrella is too close and would die too. With no other choices remaiing, Parasoul is forced to wish that Umbrella never become the Skullgirl... which of course came at the cost of Parasoul taking her place.
As Parasoul takes Umbrella home, she resolves to start Umbrella's combat training... knowing that eventually, Umbrella would be using that training against her.
Miss Nadia Fortune is a Feral (read: catgirl) cat burglar (with an appropriate pechant for cat puns) who's got herself in a spot of trouble. The gang she was in, the Fishbone Gang, managed to burgle something very important from Lorenzo Medici- the life gem that kept him young. The Medicis of course didn't take that lying down and hunted down and slaughtered the gang. Ms. Fortune lived because she swallowed the Life Gem to hide it. This didn't stop the Medici hit squad from tearing her into pieces and dropping the bits in the ocean... it just meant that when they did that, she survived, gaining the ability to heal from the most grievous of injuries and use her own body parts in combat. She wants to wish her friends back with the Skull Heart.
First, though, she stops some Medici goons from picking on her Dagonian (fishgirl) friend Minette, freaking them into running away when she bites them with her own severed head. On her way to find the Skull Heart, she fights Painwheel and Peacock before meeting Valentine, who is very interested in her body. But more in a research way than the sexual way. Ms. Fortune beats her in a fight, but Valentine is satisfied with some blood to examine, and leaves when Double calls her back to the Grand Cathedral.
Ms. Fortune then fights Parasoul before proceeding to have the customary Valentine and Double fight, followed by the Final Boss Marie. But after Marie is defeated, she has a change of heart, and decides not to throw her new life away by wishing on the Skull Heart. Instead, she destroys it and returns to Little Innsmouth... where she finds that Minette has been kidnapped. And the Adventure Continues!
Painwheel was once an ordinary schoolgirl and Filia's friend, before being kidnapped by Anti-Skullgirl Lab 0 and turned into Painwheel, an Anti-Skullgirl weapon who can sense the Skullgirl's location due to being made with Skullgirl blood, shoot spikes out from her body, and has a massive blade attached to her spine. In case this wasn't bad enough, Lab 0's head Brain Drain also mind-controls her.
She sets out to destroy the Skullgirl, fighting Peacock and Ms. Fortune before finding Valentine, who helped to turn Carol into Painwheel. Painwheel naturally wants Valentine dead, but Valentine invokes her mind control to get her to go away, forcing Painwheel to fight a copy of herself that for once isn't Double, but a representation of her programming. Valentine is surprised to see her resist the control, and tells her to meet both Valentine herself and the Skullgirl at the Grand Cathedral.
On her way there, Painwheel fights Filia, and after beating Valentine and Double, Valentine directs her to the Skullgirl. Painwheel refuses to be tempted by the Skull Heart and destroys it but becomes the next Skullgirl anyway due to her Skullgirl blood after her parents reject her.
Valentine has appeared as an antagonist on the Skullgirl's side in all previous stories, but there's more to her. She was once part of an elite Anti-Skullgirl team known as the Last Hope, but the new Skullgirl killed all of them but her, with Valentine only being spared due to potential usefulness. Now she serves the Skullgirl... but with ulterior motives.
Marie orders her to take out Lab 8, and she does so, fighting Peacock and Parasoul while she's at it. She then meets Painwheel, who unsurprisingly is really interested in killing her. After Valentine wins, she's disappointed in Painwheel's performance and points her to the Grand Cathedral if she wants a rematch. She then has a busy day fighting Cerebella, Ms. Fortune, and Filia before Painwheel finally shows up for that rematch. Double confronts Painwheel, but Valentine finally drops the act and turns on Double.
Valentine then descends into the catacombs to fight Marie, finally getting her revenge for the deaths of her teammates. After Marie is defeated, she cuts to the chase with the Skull Heart and just straight-up wishes to become the Skullgirl, as Painwheel enters the catacombs...
The mysterious Double is a shapeshifting monster who serves the Skull Heart faithfully, unlike Valentine. But that doesn't mean she's without depths or ulterior motives; note that she serves the Skull Heart, not the Skullgirl. She gave the Skull Heart to Marie, and now that Marie has proven to have a will strong enough to resist doing the normal Skullgirl thing and causing the apocalypse, Double's having second thoughts about that choice.
Double first fights Filia and Cerebella before returning for new orders, but when those orders turn out to be genuinely merciful- to attack Lab 8 but spare Peacock- Double outright ignores her orders and fights Peacock anyway, followed by Parasoul, Ms. Fortune, Painwheel, and Valentine. She then tells her mysterious master that she has gathered seven Skullgirl candidates and then proceeds to pull You Have Outlived Your Usefulness on Marie. Afterwards, she prepares to absorb all her rejected candidates...
... and cut to Venus and Aeon, who are apparently Skullgirls nerds and have been playing the game. They're called to dinner by their mother, but before they go, Aeon breaks the fourth wall to tell the audience that none of the previous story modes are canon; a true canon story has yet to be implemented.
DLC Season 1
Squigly is a cute zombie girl with a tragic story; she was originally Sienna Contiello, from a family of opera singers, before she and her family were massacred by the Medicis after a mysterious stranger (read: Double) gave her mother Selene the Skull Heart. Selene wished for her family back, but the Skull Heart is a Jackass Genie and so they were revived as her undead minions as she became the Skullgirl. Squigly only escaped her control due to the interference of the snakelike Parasite Leviathan. Now 14 years later, the new Skullgirl's necromantic powers have revived her again.
Big Band, real name Benjamin Birdland, was once a honest cop in New Meridian. Unfortunately, he was the only honest cop on his squad, and this eventually got him beaten up and left for dead. Dr. Avian then rebuilt him as a musical cyborg, and now that he knows there's a new Skullgirl, he's looking for her.
He starts his search in the River King casino, where Cerebella spots him. Given that he is a 7'7 cyborg in a trench coat, she naturally doesn't believe that he isn't there to cause trouble and picks a fight- though she probably wasn't expecting to get clobbered by a saxophone. While he wins the fight, he loses something more important- Dr. Avian. The casino was a trap; He'd show up, Cerebella would challenge him and keep him busy, and meanwhile Double and Valentine would attack Lab 8.
Big Band rushes back, but Double does her Mirror Match shtick to delay him further. By the time he gets back, Lab 8 is trashed and Dr. Avian is dead. He finds one of Valentine's scalpels in the ruins, and decides to go look up where they come from. But before that, he has to deal with Peacock, who's pissed off mightily about the attack and raring to go fight the Skullgirl. Big Band isn't keen about her going out on her own, but even after beating her in a fight, he can't stop her.
Big Band goes after her, but his search is complicated by Ms. Fortune mistaking him for a Medici (ironic, considering Cerebella earlier) and picking a fight. Again. Big Band has something of a PR problem. After Ms. Fortune gets beaten up, Irvin arrives to clarify that Big Band is friendly; he was once Irvin's partner. Irvin apologizes for Ms. Fortune and points Big Band downtown.
Luckily for him, there is some very obvious Skullgirl activity indeed- Marie and her skeletons are holding up a fallen building. Unfortunately, Painwheel is also there, and Brain Drain doesn't care that killing Marie right then would lead to the tower falling and hundreds of civilians dying. Big Band is forced to fight Painwheel when Brain Drain commands her to attack anyway. The good news is that Painwheel is eventually able to resist Brain Drain, and Big Band directs her to Lab 8's survivors, who can help her with the 'mind control' problem.
He then goes for the Grand Cathedral, to find that Marie has already defeated Peacock, and Valentine and Double fight Big Band. When he confronts Marie, he finds that Peacock is a bit beaten up but not dead, and they fight Marie together. Big Band then destroys the Skull Heart, though it taunts him with its inevitable return.
Big Band monologues that he isn't dumb enough to consider this a happy ending... but the Skullgirl's gone, Painwheel is safe and somewhere she can heal, and there was only one death. Eh, good enough!
Fukua is a bit of an odd duck. She is a clone of Filia with two souls- one a close combatant, one a ranged assassin- working together to fight. On the other hand, she may not even exist. This story is Filia's nightmare, as Fukua wants to become the real Filia, and not just a Palette Swap with a misspelled name.
Filia lies in bed one night, dreaming of a green-haired clone of herself: Fukua. Fukua fights the entire cast in the following order: Robo-Fortune, Beowulf, Eliza, Big Band, Squigly, Marie, Double, Valentine, Painwheel, Ms. Fortune, Parasoul, Peacock, and Cerebella. But wait- Marie was fought right in the middle, so who's the final boss this time? Turns out, it's Filia herself, buffed enormously in her own mind.
When Fukua defeats even this stronger Filia, she puts on a Filia mask and declares her to be the new Filia- at which point the real Filia wakes up. But when she goes to wash up, the bathroom mirror shows her reflection change to a smiling Fukua...
Eliza is a celebrity diva who runs the Bastet's Den lounge and generally stays out of the Skullgirl ruckus. Except that's not true; she's really an aeons-old Parasite/Vampire/Mummy/Catgirl abomination who caused basically everything bad in the entire game. She really does want to be a diva nowadays (it's an easy way to get attention), but the Medici have blackmail on her; they've noticed that she was embezzling blood from her charity blood drives- the blood serving to feed her parasite Sekhmet, who takes the form of her skeleton. Since she can fight with the haemokinesis Sekhmet grants her, the Medici send her off to fight things for them.
This is a terrible idea, because Eliza is not their pet attack abomination. She's older than them, scarier than them... and they've just let her loose.
Once, Beowulf was a famous wrestler, with his most famous feats being killing the Gigan warrior Grendel in single combat, followed by killing his mother when she went crazy and started attacking the crowd. But that was years ago, and now he's stuck playing a bit part in Annie, Girl of the Stars. When a new Skullgirl appears, he sees a chance to regain his fame and save the world by defeating her.
Beowulf gets a rather rude heads-up about the Skullgirl's appearance when Grendel's severed arm, which he keeps as a weapon and a memento, comes to life and starts attacking him. He goes to the Grand Cathedral to get it exorcised by Sister Agatha (Double in disguise). After she stops it from trying to murder him, he declares he's going to defeat the Skullgirl... which gives Double an idea.
Beowulf then begins his comeback tour with a fight with Cerebella. She's a big Beowulf fan, and absolutely delighted to re-introduce him to the ring, even if it does mean getting beaten up. Beowulf then goes to meet his manager, Zane, who has more fights scheduled. None of them are the Skullgirl, though; Zane instead has him cleaning up the streets by fighting some of the freaks that are running around scaring everyone.
And if you started to smell a rat when he started fighting other cast members and the "camera people" were Yu-Wan and Minette (who declare that they're just there to serve food), congratulations, you're smarter than Beowulf. He defeats Painwheel and Valentine (who does not respect him one bit and doubts he actually beat Grendel) before Annie shows up and outright tells him that something shady is going on, but he doesn't believe her and doesn't listen to her warnings not to go after the Skullgirl because fighting her isn't fake like pro-wrestling ("Hey! Wrestling is real!").
He is suspicious enough to talk to Zane about it, but Zane brushes off his concerns and then sets him up for a fight with Big Band. Big Band doesn't really want to fight, but he also called Beowulf "washed-up", so a fight it is. Beowulf celebrates his victory, but Big Band realizes that this time, it was Beowulf distracting him while Lab 8 was attacked, and quickly leaves, much to Beowulf's confusion.
Beowulf later confronts Zane again, but Zane defers his questions by promising an epic match-up... which turns out to be a Mirror Match against Double-as-Beowulf. Doublewulf tries to break Beowulf by revealing the Awful Truth that his match against Grendel was a total fraud; the Renoirs and the Medicis had teamed up to rig the match and had drugged Grendel previously.
Of course, Beowulf being Beowulf, this doesn't deter him. Instead, he confesses to the audience that his win against Grendel was faked, and promises once again to defeat the Skullgirl- this time for both them and Grendel. And he is not going to be delayed or distracted again. Which leads Zane to reveal that he'd been Double all along and eat Beowulf, transporting him to Gehenna.
Beowulf is ready to beat up Double from the inside to get out, but before he can do more than fight Double once, Annie comes in to save him and the other fighters Double ate. She scolds him for not listening to her, but he's still not deterred. He uses Grendel's arm to lead him to Marie and fights her. Afterwards, Grendel's spirit returns and animates his arm, giving Beowulf the chance to do their match right. This time, Beowulf wins fair and square, smashes the Skull Heart, and gives Grendel's arm a proper burial.
He then finally puts two and two together and realizes that this Annie and the Annie he knew in his early days are the same person. And while she thought he was an idiot at first, she came to respect him after he defeated the Skullgirl. She then decides to cancel her show and start a new one- this time with her and Beowulf as partners.
Gotta love a happy ending in Skullgirls.
Brain Drain can be a petty asshole sometimes. Like the time Valentine suggested using Ms. Fortune to help defeat the Skullgirl and he built a gynoid based on Ms. Fortune just to prove that machines were better than organics. This attempt would probably have gone better if he'd included a few less internet memes into the robot's programming, but Robo-Fortune is nonetheless a powerful fighter- and better still, it's her story mode so she gets plot armor!
DLC Season 2
Everyone knows Annie of the Stars. She's a folk hero from centuries back, and now she has a TV show based on the stories surrounding her. Annie, Girl of the Stars uses a mix of live-action and animation, and is such a Long Runner that they've had to use multiple actresses to play the part of the young heroine- long enough that the first of these actresses, Florence, is grown up and a producer. That's what everyone knows, but everyone is wrong. Annie is absolutely a real person, the stories are true, and there are no multiple actresses, just Annie playing herself. Centuries ago, she obtained eternal youth thanks to her mother's wish on the Skull Heart, and now she fights the Skull Heart whenever and wherever it pops up with the power of the stars, a sword and her remote Parasite Sagan.
Finally, Umbrella gets to show Parasoul what she's made of! The younger princess of the Canopy Kingdom, Umbrella grew up never knowing her mother or father, both having been lost in the battle against the Skullgirl 7 years ago. Parasoul is the closest thing she has to a parent, but Parasoul has a hard time juggling being a ruler and a big sister. Her only real friend is the Living Weapon Hungern, a monster that is almost, but not quite, entirely unlike an umbrella (he drools way too much to keep the rain off). And, although she doesn't know it, she has a dark hidden potential that would lead her into deep trouble if she ever encountered the Skull Heart.
This story starts with Umbrella telling the readers a story, complete with her own renditions of herself and Parasoul. In this version, Parasoul and Umbrella go to the boardwalk to have fun (albeit that Parasoul is really bad at it) instead of to the prison to see crooks Parasoul put away. Their fun/sisterly arguing is interrupted when Parasoul gets a call telling her about the Skullgirl and heads off to confront her, once again telling Umbrella to stay behind.
Umbrella then bumps into Filia, and while the two girls get along well (and Umbrella thinks Samson is cool), Samson recognizes Hungern. Worried that Hungern is going to go on a rampage and devour the city just like he did before Umbrella got him, he gets Filia to fight Umbrella. Much to Samson's surprise, not only does she win, but she doesn't die from Hungern draining her life. Filia is quick to apologize, and Umbrella is quick to forgive her.
Umbrella then goes to get ice cream, but the shop's all out thanks to Peacock having the same idea. Stymied, Umbrella decides to go find Parasoul... at the other end of the city. When she tries to climb a skyscraper to find out where she is, she nearly falls, but Cerebella saves her. The Genki Girl and Cheerful Child get along great at first, but unfortunately Eliza's also there and she recognizes Umbrella as a Renoir. She stokes Cerebella's hatred of the Renoir family to get Cerebella to agree to kidnap Umbrella despite her misgivings.
After Umbrella beats them both, Eliza decides to try again and not be so careful to keep Umbrella in one piece this time, but is sucker punched and knocked out by Cerebella, who might be cajoled into kidnapping a child but wants no part in mutilating one, even if the kid is a Renoir. Cerebella then gives Umbrella some Brutal Honesty about what her parents got up to and tells her to leave, still unsatisfied since yelling at a Renoir also meant making a child cry.
While Umbrella is wandering the streets of New Meridian dejected about how badly her parents screwed up, she meets Beowulf. And after she beats him up when he tries to 'rescue' her from Hungern, he gives her probably the best advice she'd ever heard: you can't change the past, but you can learn from it, and Umbrella could do right by her parents by not making their mistakes. Now cheered up, Umbrella goes back to find Parasoul.
Umbrella finds Parasoul, but the two once again argue. Parasoul thinks that their family has lost enough and doesn't want to lose Umbrella, but Umbrella points out that Parasoul's the only family she has left, so the same argument applies to Parasoul. Umbrella's Skullgirl energy spikes, and their argument ends with an actual fight and Umbrella running away.
Both parties immediately regret the fight, but also both think it's too late to make amends. While running away from Parasoul, Umbrella runs into Painwheel, who's conflicted with her desire to reunite with Filia and her programming to fight the Skullgirl- especially since Umbrella feels remarkably Skullgirl-like. Brain Drain controls Painwheel to fight Umbrella, but after Painwheel is defeated, the resonance between her and Umbrella's shared Skullgirl blood backlashes against Brain Drain and breaks his control for good- and good riddance.
Turns out, without Brain Drain, Carol is a really nice person and encourages Umbrella to go back to Parasoul, telling her that if Parasoul really loved her sister, she would've already forgiven Umbrella for the fight. Umbrella in turn encourages Parasoul to reconnect with her family by saying that she actually looks cool as Painwheel.
It's just in time, too, as Double has separated Parasoul from most of her forces, and Marie has cornered her. Umbrella, Painwheel, and the Egrets go to save her, and Parasoul and Umbrella have a heart-to-heart talk as the Egrets and Painwheel take on Marie in the background. Umbrella finally is able to get Parasoul to accept that she too needs protected, and Umbrella is now an implemented fighter and can hold her own. The two sisters then double-team Marie, and after she's defeated, Umbrella confronts the Skull Heart.
This nearly ends like it did in Parasoul's story mode, but Hungern and Parasoul both snap her out of it. Umbrella has now had some character development and can recognize that the Skull Heart isn't her mom. She and Parasoul then destroy the Skull Heart together.
And that brings us back to the present, where we see that Umbrella has been telling the story of how she beat the Skullgirl to Filia and Painwheel, who are having a sleepover with her and Parasoul. Once Umbrella is asleep, Parasoul goes to talk to someone who's quite happy with how she and Umbrella have been growing up... her father, King Franz. He's still alive, but no longer quite human as he as a pink flame instead of a head.
Black Dahlia is a Straw Nihilist and hitwoman for the Medicis who really doesn't care for the Skull Heart; all she cares about is alcohol, cake, and violence- mostly the latter. She was rebuilt long ago by the Anti-Skullgirl Labs, but she's about as interested in fighting the Skullgirl as she is in wishing on the Skull Heart.
You know her, you love her. Marie has been the titular Skullgirl for ten years now, and all routes have ended with her dead and/or the Skullheart destroyed. That's how the story of every Skullgirl ends; with some brave hero killing the Big Bad and either doing the smart thing and smashing the Skull Heart, or the stupid thing and wishing on it, becoming the new Skullgirl that some other, wiser hero must defeat.
But Marie's story is not yet over. The Skull Heart is gone, but she still lives, the first Skullgirl in history to be completely free of its influence. And while Marie isn't completely sure what to do with her newfound freedom, she is sure of one thing: She will be the last Skullgirl.
