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Paris, 1941. A family of scientists is on the brink of discovering a powerful longevity serum when suddenly a mysterious force abducts them, leaving their young daughter, April, behind. Ten years later, April (voiced by Academy Award winner Marion Cotillard) lives alone with her talking cat, Darwin, and carries on her family’s research in secret. But she soon finds herself at the center of a shadowy and far-reaching conspiracy, and on the run from government agents, bicycle-powered dirigibles, and cyborg rat spies.
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Paris, 1941. A family of scientists is on the brink of discovering a powerful longevity serum when suddenly a mysterious force abducts them, leaving their young daughter, April, behind. Ten years later, April (voiced by Academy Award winner Marion Cotillard) Creator/MarionCotillard in French) lives alone with her talking cat, Darwin, and carries on her family’s research in secret. But she soon finds herself at the center of a shadowy and far-reaching conspiracy, and on the run from government agents, bicycle-powered dirigibles, and cyborg rat spies.
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* AlternateHistory: Where the Franco-Prussian War never happened and where a string of serial kidnappings of brilliant scientists has left the world in a state of technological stagnation.
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* AlternateHistory: Where the Franco-Prussian War UsefulNotes/FrancoPrussianWar (and therefore, neither UsefulNotes/WorldWarI nor UsefulNotes/WorldWarII) never happened and where a string of serial kidnappings of brilliant scientists has left the world in a state of technological stagnation.
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* AnimalsLackAttributes: Averted; Darwin’s genitals are visible a few times in the film.
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* {{Infodump}}: A rather lengthy worldbuilding narration of the world and how it begins to turn into AlternateHistory once a war is avoided.
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* {{Infodump}}: A rather lengthy worldbuilding narration The intro, which is a bit of a throwback to the comics and silent film serials that inspired the moview, telling the audience about the state of the world world, the AlternateHistory, and how it begins to turn into AlternateHistory once a war is avoided.the wars being fought.
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* WomenAreWiser: Chimene is more pragmatic than Rodrigue and is constantly telling him to be more gentle when interrogating and searching for April.
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* WomenAreWiser: Chimene is more pragmatic than Rodrigue and is constantly telling him to be more gentle when interrogating and searching for April. She also turns out to [[spoiler:be the one who isn't a psychotic genocidalist.]]
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* FauxAffablyEvil: When [[spoiler:Rodrigue]] drops the charade he changes from a brusque and perpetually annoyed personality to a much more theatrical one dripping with mocking niceties.
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* KarmaHoudini: Not only does [[spoiler:Pizoni]] never get any lasting comeuppance for his actions, he also takes the credit for [[spoiler: foiling the lizards and saving the scientists]] which gets him promoted to [[spoiler: head of the French Secret Service by the Emperor himself.]]
* KarmicDeath: Napoleon III is killed after he orders his minions to fire at Chimene and Rodrigue in a lab full of volatile chemicals.
* KnightTemplar: Late in the film [[spoiler:Rodrigue]] goes from a WellIntentionedExtremist to this. [[spoiler:Like Chimene, he too seeks a better world, but unlike her he doesn't want to spread plant life but rather kill off humanity and establish a new world run by him and the other lizards]].
* KarmicDeath: Napoleon III is killed after he orders his minions to fire at Chimene and Rodrigue in a lab full of volatile chemicals.
* KnightTemplar: Late in the film [[spoiler:Rodrigue]] goes from a WellIntentionedExtremist to this. [[spoiler:Like Chimene, he too seeks a better world, but unlike her he doesn't want to spread plant life but rather kill off humanity and establish a new world run by him and the other lizards]].
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* KarmaHoudini: Not only does [[spoiler:Pizoni]] never get any lasting comeuppance for his actions, he also takes the credit for [[spoiler: foiling the lizards and saving the scientists]] which gets him promoted to [[spoiler: head of the French Secret Service by the Emperor himself.]]
* KarmicDeath: Napoleon III is killed after he orders his minions to fire at Chimene and Rodrigue in a lab full of volatile chemicals.
* KarmicDeath: Napoleon III is killed after he orders his minions to fire at Chimene and Rodrigue in a lab full of volatile chemicals.
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* KnightTemplar: Late in the film [[spoiler:Rodrigue]] goes from a WellIntentionedExtremist to this. [[spoiler:Like Chimene, he too seeks a better world, but unlike her he doesn't want to spread plant life but rather kill off humanity and establish a new world run by him and the other lizards]].
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* KnightTemplar: Late in the film [[spoiler:Rodrigue]] goes from a WellIntentionedExtremist to this. [[spoiler:Like Chimene, he too seeks a better world, but unlike her he doesn't want to spread plant life but rather kill off humanity and establish a new world run by him and the other lizards]].
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* BigBadWannabe: The French Government. Napoleon III kicks off the plot with his lust for war but he's killed rather quickly because of his stupidity. The government begins kidnapping scientists to build up their army and hunts down April's family at the beginning and middle of the film. Though they're a constant threat, they ultimately pale in comparison to the real villains of the setting and they fall out of focus for the third act.
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* FreudianExcuse: Since one of his earliest experiences with humans involved them trying to kill him (and left him with a permanent facial scar), [[spoiler:Rodrigue]] grew to hate them
* KickTheDog: Downplayed. To demonstrate the serum, [[spoiler:Rodrigue]] shoots [[spoiler:Darwin]] in the chest. He did so with the knowledge that [[spoiler:Darwin]] would be fine, but even then it's still a painful act that [[spoiler:Chimene]] criticizes as unnecessary.
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* ParentalSubstitute: with the loss of her parents and Grandfather, Darwin acts as a parental figure for 10 years of April's life.
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* ParentalSubstitute: with With the loss of her parents and Grandfather, grandfather, Darwin acts as a parental figure for 10 years of April's life.
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* ParentalSubstitute: with the loss of her parents and Grandfather, Darwin acts as a parental figure for 10 years of April's life.
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* KidnappedScientist: One of the reasons why scientific progress has been retarded is renowned scientists such as Einstein and Fermi mysteriously disappear just as they are becoming famous. The scientists are being held prisoner by the {{Big Bad}}s to work on their secret project.
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* PostApocalypticGasMask: It is established that is not safe to walk the streets on heavy pollution days without a respirator mask. However, only the wealthiest in society can afford them. This does afford Pops an excellent disguise, as no one looks twice at an elderly man walking around in a gas mask.
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* PostApocalypticGasMask: It is established that it is not safe to walk the streets on heavy pollution days without a respirator mask. However, only the wealthiest in society can afford them. This does afford Pops an excellent disguise, as no one looks twice at an elderly man walking around in a gas mask.
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* BalconyEscape: April's parents attempt to escape from the secret police by climbing off the balcony and clambering across the outside of the cable car.
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* EvilutionaryBilogist: Napoleon III wanted Dr. Franklin to create him an unstoppable army of SuperSoldier animals with which to crush Prussia. He was not happy when Dr. Franklin came up with {{Uplifted Animal}}s instead.
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* EvilutionaryBilogist: EvilutionaryBiologist: Napoleon III wanted Dr. Franklin to create him an unstoppable army of SuperSoldier animals with which to crush Prussia. He was not happy when Dr. Franklin came up with {{Uplifted Animal}}s instead.
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* EviloutionaryBilogist: Napoleon III wanted Dr. Franklin to create him an unstoppable of SuperSoldier animals with which to crush Prussia. He was not happy when Dr, Franklin came up with {{Uplifted Animal}}s instead.
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* EviloutionaryBilogist: EvilutionaryBilogist: Napoleon III wanted Dr. Franklin to create him an unstoppable army of SuperSoldier animals with which to crush Prussia. He was not happy when Dr, Dr. Franklin came up with {{Uplifted Animal}}s instead.
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* EviloutionaryBilogist: Napoleon III wanted Dr. Franklin to create him an unstoppable of SuperSoldier animals with which to crush Prussia. He was not happy when Dr, Franklin came up with {{Uplifted Animal}}s instead.
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* ReassignedToAntarctica: After losing the Franklins, Pizoni is demoted from inspector to agent, and told that from now on he is not allowed anywhere scientists. He is restricted to arresting tramps and vegetable thieves.
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* ReassignedToAntarctica: After losing the Franklins, Pizoni is demoted from inspector to agent, and told that from now on he is not allowed anywhere near scientists. He is restricted to arresting tramps and vegetable thieves.
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* XRaySparks: Happens every time someone is electrocuted. First seen with the gendarmes and then April's parents on the cable car, and later with Julius in Pops' mansion, and then various people in Rodrigue and Chimène's base.
