Characters / Small Saga

Main Protagonists

Verm

• Abdicate the Throne: At the end of the game, with King James dead and no direct successors stepping up out of fear of the revolutionaries, it seems that Verm could claim the throne for himself. Instead, he sticks the Titan Cleaver into the throne, declares that it's time for a change of governance, and leaves.
• Ambiguously Absent Parent: At the start of the game, Verm's brother Lance mentions "dad's letter" and talks about him in the past tense, but nothing more is shared about the family - whether either of their parents are estranged or alive at all, if Verm's gone home or seen any other family since losing his tail, etc. The developer says that some things are best left up to the player; any answers they can come up with will be more interesting than a bit of Word of God.
• And This Is for...: During the fight with the Yellow God, Verm will say different things when he uses Berserk. In the first phase he says "This is for my tail," and in the second he says "This is for my brother."
• Attack the Tail: In the prologue Verm's tail is pinned by a Mouse Trap and he eventually has to make a Life-or-Limb Decision and hacks through it with a broken sword. This is portrayed as a combat situation, with each blow taking a substantial chunk of his health.
• Beware My Stinger Tail: The iron tail prosthesis Gwen makes him is barbed at the end. He doesn't actually use it as a weapon, but it makes him substantially faster and more agile and the look of it has Leo calling him a "demon".
• BFS: The human-sized pocketknife he wields is considered one of these, to the point where many of his opponents are baffled that he can even swing it at all. Its size is a key factor in the battles against Blademaster Leo, whose rapier-like needle blade shatters under a few good blows from Verm's weapon.
• Blood Knight: Downplayed. If threatening presences like Tiger the cat are willing to let him pass he won't offer violence. His beef is with the Yellow God. However, he's quite eager to fight if they don't yield and tends to yell rather psychotic phrases while Berserking.
• Broken Pedestal: Played With with regards to Blademaster Leo, who he absolutely idolizes in the prologue. Discussing this with Gwen, she believes that meeting Leo showed Verm that he wasn't all that, but Verm does still think highly of him.
• Cathartic Chores: During the Time Skip Verm is fished out of the sewer by a shrew who saved him to make him into a farm worker. Verm, feeling adrift and that vengeance was hollow, dedicated himself to mushroom farming with the same single-minded intensity he'd put to fighting.
• Chaste Hero: While exploring Murida with Gwen, Verm has no idea that the occasion is supposed to be a date. During said date, he can have a conversation in which he says that he's never had the time to really consider and is not really sure what kind of person he's interested in, if anyone. This is reinforced when he looks at the smutty drawings in the old contraband storage room under Sky Garden (which contain images of both male and female rats), with his only response being "Huh."
• Cynicism Catalyst: In the first dungeon, Verm is rude to enemies but is playful talking to his brother and jumps enthusiastically in celebration upon winning fights. The encounter with the Yellow God hardens him a great deal. One signifier is his victory animation - now he folds his arms and spits on the ground upon defeating enemies. As he learns to trust and rely on his party and gains a bit more wisdom, this changes again to folding his arms and nodding.
• The Drifter: At the end of the game he wanders off without a plan of where to go or what he wants to do, though he mentions maybe returning to the place where he and Lance came from.
• Expy: Verm is a mouse version of Guts in a less harsh and dire world. From the cloak and handwraps and BFS, to making a Life-or-Limb Decision and severing a part of his body with a broken sword, losing an eye, getting a prosthetic that should take him a long time to become accustomed to but doesn't, going against a force far far above him, being rather terse except in battle where he can show Blood Knight aspects... One of his lines is "As long as I was swinging my sword at a foe, I didn't have to think" which is an extremely Guts thought. There's also a shot of him riding an owl while holding the Titan Cleaver that's a clear visual reference to Guts riding on Zodd's back. Verm's dark period is not nearly as pronounced or sinister as the Black Swordsman's of course, but he does learn the same kind of lesson about doing better with friends and allies and contemplates giving up on Revenge, but Verm is more easily able to manage it as he has no analogue to the Berserker Armor and humans may be Cthulu but aren't Griffith.
• Eyepatch After Time Skip: No eyepatch, but after the Time Skip he's more scarred and one eye is missing, the eyelid always closed. Again, this is just like Guts. It does seem like he got this either in his duel with Leo or upon falling into the sewer afterwards, not in some off-screen adventures, as he spent the entire season peacefully farming mushrooms.
• Giant's Knife; Human's Greatsword: His "Titan Cleaver" is just a normal human pocketknife.
• Handicapped Badass: He loses his tail in the prologue, which a few characters note would probably throw off his balance, but is more than capable in combat despite this. When he gets a prosthetic tail, he becomes much more agile.
• Instant Expert: When Gwen gives him a prosthetic tail, she warms him that it might take him weeks to get used to it. He acclimates almost immediately.
• Jerk with a Heart of Gold: He's rude to prospective enemies but overall excitable and idealistic in the prologue, which changes afterwards. The player has occasional dialogue choices that can control the degree of "jerk" at times, but even so Verm in the early game really only cares about his quest for revenge and is quite abrasive and dismissive of other concerns. However he can't bring himself to actually leave an ally in the lurch; when he starts to walk out on Siobhan early on he can't and tells them "You're pathetic. I'll help you." Gradually he mellows out.
• Life-or-Limb Decision: Early in the game, Verm's tail is pinned by a mousetrap. Lance tries to heft the bar up to let Verm pull free but can't manage. Lance then goes to distract the approaching Yellow God and is killed. Verm tries to attack the mousetrap but only manages to break his sword, and finally uses his broken sword to sever his tail and escape.
• Nominal Hero: Much of the time if he does something good for others it's because it will clearly also benefit him somehow, or because Siobhan talks him into it, or because he feels like walking away is the same as him running away from danger. Post-timeskip he's less of a jerk and thinks more of his friends and allies but he's pretty indifferent to the big picture and the future. Leo's asking And Then What? if Verm manages to kill a God doesn't move Verm, but seeing himself reflected in the Yellow God's eyes and understanding that he's changing for the worse does.
• No One Gets Left Behind: After the events of the opening, Verm has several points where he tries to leave friends or allies to fend for themselves, only to get a splitting headache. He seems to flash back to Lance, saying "I'm not running away!" and then he always commits regardless of the risk.
• Rebel Relaxation: He's often seen leaning his back against a wall, arms folded, one knee bent.
• Sibling Yin-Yang: Verm is much ruder than his brother Lance.
• Spiteful Spit: For much of the game his victory animation on winning a fight is to fold his arms and spit on the ground.
• Unscrupulous Hero: Verm's really only interested in Revenge for most of the post-prologue story and is quite dismissive of everything and everyone that can't help him in that endeavor. He does get some Character Development and mellows out a bit.
• Vengeance Feels Empty: When he faces Leo, Leo tells him that he fights for nothing and his attacks lack weight. Throughout the Time Skip Verm ponders this and even attempts to give up on revenge as impossible, but when he enters the gladiatorial arena to help Bruce he takes his goal up again. At the end of the game, Verm ends up showing mercy to the Yellow God he fights, and lets him flee. As he tells Siobhan later, he's fully aware that any human could wear the Yellow robes and it was impossible to truly avenge his brother, but was still using that quest for vengeance as a way to give himself purpose until he saw his reflection in the God's eyes and realized he'd become a ghost of his former self.

Siobhan

• Ambiguous Gender Identity: In the early game, Siobhan doesn't like when their Mammy calls them "young lady", but sometimes self-introduces as Lady Siobhan, and friends refer to her with feminine pronouns. After the Time Skip, Siobhan's friends use they/them pronouns for them and they tell their mother's Blademaster that they're not a girl anymore, but nothing is specified. This likely came from working under Aquilla as their assistant, because Aquilla personally considers the idea of gender distinction to be utter nonsense compared to the pursuit of scientific knowledge.
• Bad Liar: As the player you sometimes get to choose if Siobhan responds to a question by fessing up or trying to hide things. When the latter is chosen, other characters always find their attempts to lie very transparent.
• Beware the Nice Ones: Siobhan's the kindest member of the party but can do huge damage. If they really take against someone, watch out.
• Glass Cannon: Siobhan's HP is lacking, but their lighter deals out absurd amounts of damage when fully overclocked, to the point where they can single-handedly reduce endgame bosses' HP by half in a single turn.
• Morality Chain: Siobhan is convinced that Verm is Sir Verm, a heroic knight who will of course help those in need. Even if the player chooses nicer and more diplomatic dialogue options when they're available, Verm is quite callous early in the story and initially he doesn't want Siobhan following him, but becomes resigned to it. Repeatedly Siobhan insists on taking on causes and convinces Verm that they'll benefit him as well, and acts like his hostility is a joke. A conversation with Aquila suggests that they are aware of how indifferent Verm actually is to things that aren't going to help in his quest for vengeance, but that's post Time Skip.
• Never Learned to Read: Siobhan can certainly read rodent writing but hasn't learned God writing, as they mention while showing Verm their little hoard of God items. In Aquila's lab, which is half taken up by human books, they mention this again and wonder if Aquila will teach them.
• Quivering Eyes: Gets this after being exiled from Cranbaile.
• Rebellious Princess: Their "Mammy" is Gilda, the leader of Cranbaile, a tiny kingdom that sticks to the Old Ways of nonaggression with the Gods and their Titans despite their own citizens being endangered by a housecat. Siobhan is eager to fight back and then to join Verm on his quest despite his reluctance, but their mother is in such denial that she condemns Verm as a corrupter and kidnapper.
• Robe and Wizard Hat: After the Time Skip, Siobhan wears a pointy, wide-brimmed hat that looks like a classic witch's hat. The hat is actually traditionally worn by males, showing their increased comfort with their gender nonconformity.
• The Smart Guy: Siobhan is almost invariably the one to fill the other characters in on anything technical or scientific. Aquilla even takes notice of this and, despite their high standards and overall issues with the rest of Muria for their experiments, is completely genuine about allowing Siobhan firstly to become their assistant, then to inherit the title of Plaguemaster of Muria after their passing.
• Smart People Wear Glasses: Real moles have very poor eyesight, but most of the ones in the game don't wear glasses. Siobhan does, though, and they signify the above trait.
• Unusual Weapon User: Siobhan wields a souped-up cigarette lighter that's been modified to send out blasts of flame.

Bruce

• The Bard: He mostly contributes to combat by playing songs (through a flute made from a hollowed-out pencil) that heal and empower the other characters.
• Booze-Based Buff: One of his special skills quintuples his damage when he's tipsy.
• Dreadful Musician: His post-timeskip quest starts with Anton and Gwen complaining that he's not putting his heart into his music and his high notes are shrill, leading him to jam out with two other musicians to try and get his groove back, culminating in Hey, Let's Put on a Show.
• Everything's Louder with Bagpipes: They don't actually appear in-game but going with his Scottish-ness, bagpipe frequently appears in songs associated with Bruce.
• Instrument of Murder: His one damage-dealing attack is to thwack enemies with his pencil flute.
• Man in a Kilt: His second outfit incorporates a kilt and sporran.
• Performer Guise: He and Anton's plan to move against Clan Grey has them performing music in front of the Duke to distract him as an accomplice steals the Stone of Scuin.
• Significant Wardrobe Shift: Bruce begins wearing a blue bodysuit with matching mask because he needs to hide his identity in public due to Clan Grey slaughtering Clan Red. After Clan Grey has been ousted, he switches to a blue kilt, white shirt, and uncovered face.
• Support Party Member: He doesn't even have a way to deal damage to enemies at first, only able to play songs that heal and empower his allies. While he does eventually get an attack, it's not all that strong and his songs are going to be used far more.
• Tipped Off by the Tail: As they're approaching the Ash Tower, Anton tells Bruce to "tuck" and hide his tail. His paws, nose, and ears still show red under his costume but apparently it's the tail that will give him away as a red squirrel in gray squirrel territory. Indeed when Duke Josh become suspicious of him, he demands to see Bruce's tail.

Gwen

• Affectionate Nickname: She calls Verm "dwt" (pronounced "doot"), a Welsh word that basically means "little guy."
• Amazon Chaser: The reason that she tried to flirt with the massive and terrifying Blademaster Lamia. Her response to Bruce's incredulous statement that she'd eat Gwen alive is "Gods, I wish."
• An Arm and a Leg: Gwen is a former lab rat whose hind legs were both removed and replaced with prosthetics as part of an experiment.
• Black-and-White Morality: Gwen divides the world into good people and bad people and thinks at best, bad people should be cut out of one's life without question, and that a Freudian Excuse Is No Excuse.
• Cop Hater: None of the party members are impressed with "guards" and one of Gwen's skills is even "All Guards Are Busted". Despite never speaking to or interacting with him in any way, Gwen despises Leo and thinks he's Obviously Evil for being part of the system, with no heroic or sympathetic aspects to speak of, and doesn't understand why Verm would be ambivalent about him. If Verm spares his former hero, Gwen disparagingly says that she would have gutted him.
• Escaped from the Lab: She shook off her previous acceptance of the Old Way and left after the White Gods removed her legs and gave her new ones.
• Giant's Knife; Human's Greatsword: Her "glaive" is a scalpel.
• Handicapped Badass: Both of her legs are prosthetic, but it doesn't seem to slow her down much.
• Huge Guy, Tiny Girl: Her ex-boyfriend, sir George, is a rabbit that absolutely dwarfs her.
• Indirect Kiss: She tastes Verm's macha latte and then comments that this was an indirect kiss.
• Lab Pet: She's a white ex-lab rat and dated a white lab rabbit for a time.
• Out of the Closet, Into the Fire: She was stuck into the Clan Grey dungeon for the "crime" of being queer in a territory that doesn't tolerate it.
• Overly Long Name: Gwenllian of Neuadd-y-duwiau-gwynion, the Hall of the White Gods.
• Raised in a Lab: It's unclear whether or not she was born and grew up in the Hall of the White Gods or taken there at some point. Real animal testing usually involves procuring test subjects from elsewhere rather than breeding them but, well, these aren't real animals.
• Secret Test of Character: She could have slipped her chains at any time by just detaching her leg, but sent Bruce out to get the key to see if he would be able to do it and reliable enough to come back for her.
• Tiny Guy, Huge Girl: When matched with Verm, she's more than twice his size. However, the people she's romantically attracted to are taller than her.
• Unstoppable Rage: When confronting her old boyfriend/mentor George over the whereabouts of their missing friends and family from White Hall, Gwen initially keeps demanding answers from him between trading attacks, but in the face of his persistent silence and disapproval over her choice to defy the Gods who "care" for them, Gwen eventually unleashes her own version of Verm's Berserk state, only hers lasts for a whopping 98 turns, more than enough to finish the fight with, especially since it occurs when George's health is getting low.

Allies

Lance (Prologue Spoilers)

Verm's older brother, who has a plan to break into Heaven and make off with a godly hoard.

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• Curb-Stomp Battle: His attacks do absolutely no damage to the Yellow God, and when it turns its attention to him it immediately grabs him in its hand and crushes him to death, dealing hundreds of times his maximum health in damage.
• Heroic Sacrifice: When Verm's tail is caught in a mousetrap and the Yellow God approaches, Lance can't get him free and races up some shelves to the God's face level, determined to distract it before it can catch Verm.
• Interface Spoiler: Masterfully subverted. Lance has an entire skill tree that is just as fleshed out as any later party members', leading the player to think they might survive before their very unambiguous...
• Plot-Triggering Death: His death to the Yellow God is what kicks off Verm's quest for vengeance.
• Sibling Yin-Yang: He's more friendly and polite, including to strangers, than his little brother Verm.

Anton

Bruce's boyfriend, a hamster.

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• The Cavalry: Completing Bruce's endgame sidequest leads to Anton joining the revolution and fending off some knights at a key moment.
• Distressed Dude: After Anton and Bruce are both apprehended by Clan Grey, he's kept in a separate cell, and rescuing him before Lamia decides to eat him becomes one of Bruce's main goals.

Clan Grey

In General

• Eagle Land: Type 2, patriotic American-accented colonizers from across the sea who exterminated the native red squirrels to take over their territory. However, most of the vileness comes from their leadership, with their actual citizens portrayed as victims of a tyrannical regime.
• Politically Incorrect Villain: Aside from their Fantastic Racism to everyone who isn't a grey squirrel, they're also homophobic.
• Salt the Earth: Clan Grey did this to the original Sky Garden in order to take control, burning its tree into an ashen pillar, killing the Cailleach, and slaying all of Clan Red save for Bruce.

Duke Josh

The leader of Clan Grey.

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• Evil Is Not a Toy: He commands the loyalty of Lamia, a rodent-eating stoat. She'll keep Sky Garden safe against all enemies if in return she's fed every time she's hungry, so he lets her use his dungeons as her personal larder. Lamia thinks he's a bit too casual about this. After he's overthrown, Lamia comes to him saying she's hungry, and since he has no one else to offer her she makes him her next meal.
• Fed to the Beast: He tells Lamia to eat people he's found particularly aggravating.
• Good Smoking, Evil Smoking: He regularly takes hits from a (human) vape pen, signifying his crass, obnoxious personality.
• Karmic Death: Duke Josh leaps from Sky Garden to flee after being defeated by the party, but wounds his ankle on the landing. The only subordinate that comes to his call is Blademaster Lamia, who's hungry... and with no one else he can give her to devour, she turns on him.

Lamia

Duke Josh's Blademaster, known as "the Peerless".

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• Ax-Crazy: Blademaster Lamia is a stoat that threatens to eat her own allies, but is tolerated by Clan Grey because their Duke made a deal with her. She's a peerless warrior and as long as she's fed whenever she gets hungry she will take on all threats. Mostly she eats the Duke's enemies and prisoners. In the boss fight against her, she outright cannibalizes her subordinate to heal herself, and abandons her skewer weapon in favor of biting and clawing at the party.
• Comfort Food: Lamia complains of feeling great loneliness without other stoats; she's peerless in that she has no real allies, either. Clan Grey despises her, and she feels a void that she fills by devouring rodents.
• Double Meaning: In the fight against her, she explains that she's not just "the Peerless" because of her remarkable combat skills; she's also the only stoat that did not flee to the Outwoods, leaving her without literal peers.
• Giant's Knife; Human's Greatsword: She wields a barbecue skewer as a lance.
• Neck Lift: Blademaster Lamia does this when Hazel the janitor chooses not to squeal on the party after they pull a jailbreak to save Bruce and Gwen. She also nearly eats him, but is stopped by a subordinate.
• Uncertain Doom: The party beat her to hell and kick her out the top of the Sky Garden tree, but she shows up finish off Duke Josh after he likewise leaps from the top to escape the group. Despite being bloodied she goes after him with no trouble, and it's unclear whether she succumbed to her injuries after her meal or recovered and fled. The party have other concerns to deal with regardless, and she's no further trouble.
• Wicked Weasel: She follows the terms of her arrangement with Duke Josh to the letter, but isn't above extracurricular snacking.
• Wham Shot: If you thought the game had eased up on the red-in-tooth-and-claw since the introduction, time to think again: Lamia eats one of her subordinates right in front of you.

Rodentia Royal Court

Sir Alex

One of the king's knights, who has a rivalry with Blademaster Leo. He was Blademaster in Needle Knight.

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• Dirty Coward: He's rather underhanded and cowardly, exemplified in an endgame sidequest in which he tries to hire Verm's party to kill his former subordinates Rosalie and Maisie for insubordination, after he used Maisie as a Human Shield to save himself and Rosalie attacked him in retribution.
• Drowning My Sorrows: He's found lying in the alley outside of the Twisted Tails tavern drunk on some concoction that smells like vinegar after Rosalie and Maisie turn on him.
• General Failure: Regardless of his swordsmanship, Sir Alex is an abysmal tactician. When he's given command of the kingdom's forces to quell the riots in the final act, he's so deranged by his need to show up Leo, who he refers to as "that blond twink", that he marshals the entire guard to the palace's front door — including the guards who were patrolling the palace, leaving the King almost defenseless.
• Green-Eyed Monster: He's bitterly jealous of Sir Leo for taking his position as the kingdom's Blademaster.
• Karmic Death: In the game's Playable Epilogue, according to Steve, Sir Alex the Sterling Knight managed to survive the revolution by fleeing the kingdom, only to try to swim for safety and sink like a lump of silver.
• Obfuscating Stupidity: Bree and Stilton are fomenting revolution in the Twisted Tails tavern, and while Sir Alex is soused he hears enough to take it to the King.

Rosalie and Maisie

The scissor-sisters, one veteran and one promising new recruit who work for Sir Alex.

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• Goldfish Poop Gang: The siblings encounter Verm's party a few times throughout the story. Rosalie is the more competent of the two, but neither of them are a match for the party.
• Find the Cure!: In the late game after Sir Alex used Maisie as a Human Shield, she was bitten by the Doom of Solhill. This didn't immediately kill her but had her weakening steadily. Verm and his party can help by fighting the Doom themselves and collecting some venom while Rosalie gathers the other ingredients to make a cure.
• Heel–Face Turn: A late-game sidequest to save Maisie from snake venom convinces Rosalie to join Verm's side in the rebellion. It's not a Magic Antidote, so while Maisie is game too she's still too ill to participate in the revolt herself.
• Human Shield: Maisie ends up used as one by Sir Alex to save himself from a Titan snake, which outrages Rosalie and leads to them both leaving Alex's service.
• Shear Menace: Each of them wield one half of a pair of scissors as a sword.
• Those Two Guys: Rosalie has a role in Needle Knight where she's the most professional of the band of Thimble Guards sent to Solhil. In the game itself the sisters are rarely seen apart, and always comment on trying to find the other or where the other is when they are apart.

Sir Leo

The Royal Blademaster, an honorable and legendary knight who was the protagonist of Needle Knight.

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• Animal Motifs: He refers to himself as a golden lion a couple of times, referring to both his name and his yellow-tinged fur.
• Begin with a Finisher: The second fight against him has him use Fleche Fatale on his first turn, the same One-Hit Kill move he used on Verm in his first duel. However, Verm can either attack and fatally wound him before he gets the chance, or use Nimblefoot to dodge the attack entirely.
• Embarrassing Nickname: In Needle Knight he starts with the nickname "Lemon" after embarrassing himself in a Noodle Incident.
• Field of Blades: The final battle against him takes place in one of these, with his numerous spare needles scattered around the throne room.
• Hero of Another Story: Literally in that Needle Knight showed him proving himself against a Vulpes that was picking off the residents of Solhil. The webcomic stopped before actually showing this fight, but there's an in-universe song about it and statues have been raised in his honor. A book in Vinium also describes him fighting several duels to become Blademaster, taking the title from the more cowardly Sir Alex. Then there's his off-screen fight with Diego and the Thimble Guard after he murders his king, which leaves Diego's hammer and all the guards' swords strewn across the throne room.
• Hopeless Boss Fight: The first Duel Boss against Blademaster Leo has him parry everything Verm throws at him. Even after Verm manages to break Leo's weapon, he still follows up with an attack with his broken weapon that deals several times Verm's health.
• Legendary in the Sequel: He was the protagonist of the Needle Knight prequel comic, where he started as a bumbling new recruit.
• Luke, I Am Your Father: Downplayed. Sparing him in the rematch by breaking all his needles will have someone comment that he left town with his daughter- his daughter that the general public was unaware of. It's implied his daughter is Virgo, a young mouse who complains of her dad working late and can help Verm read a treasure map.
• Master Swordsman: Regarded by many to be the greatest living swordsman in Rodentia, with fighting strength equal to an entire battalion.
• Promptless Branching Point: In the endgame, whether or not Leo survives is dependent on the duel against him. If Verm defeats him, Leo dies; if Verm stalls long enough for all of Leo's spare needles to break, Leo surrenders and lives.
• Rebel Relaxation: Unlike the Loremaster, who stands at the King's side, Leo is usually sitting and lounging with one of his needle swords stuck into the floor. When Verm enters the throne room after Leo kills the king, Leo's sitting like that on the throne.
• Rogue Protagonist: Sir Leo, of Needle Knight, is now the direct underling of the king that forbids conflict with the Gods, which is Verm's primary motivation. Near the end of the game, Leo duels Verm to the death out of concern that Verm's crusade of vengeance will lead to a war with the Gods that will doom everyone.
• Sewing Needle Sword: What he's known for, using them like rapiers.
• To Be Lawful or Good: Upon realizing that King James is willing to sacrifice his own subjects to keep himself in power, Needle Knight Leo has a brief monologue about what it really means to be honorable before turning traitor and stabbing the King in the back.
• Unwitting Instigator of Doom: His slaying of the fox that was menacing Solhill did save them from the beast, but its rotting corpse spread disease and attracted far more dangerous predators, eventually leading to the death of everyone in Solhill.
• Wrecked Weapon: The Duel Boss fights against Leo will have Leo parry Verm's attacks, but his sewing needles can only take so many hits before breaking. The first time, he's still capable of attacking and one-shotting Verm with a broken needle. In the second fight against him, Leo has several spare needles around the arena as replacements if his weapon gets broken. However, once all of them run out, the fight ends with Verm victorious and Leo being spared.
• Zodiac Motifs: Like the sign he's named after, Leo is a charismatic, confident knight, with a penchant for showsmanship. He is also courageous enough to betray his own king, but only after it becomes clear he values nothing for the lives of his own people. It's also implied this is a family tradition- the girl implied to be his daughter in the ending where you spare him is named Virgo.

Sir Diego

Leo's closest companion, the gigantic guardian of the throne room.

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• Brutish Character, Brutish Weapon: He's so huge he totes a human gavel as his weapon and often speaks with the same intimidating dramatic script as Titans.
• Expy: In appearance, though not personality, he's clearly inspired by Executioner Smough.
• Gate Guardian: While Leo stands beside the throne, Diego stands in front of the gate to the throne room. Try to enter without permission and Diego slams his hammer on the ground with enough force to knock you off your feet.
• To Be Lawful or Good: Like Leo, he has his doubts about King James as a ruler. Unlike Leo, he comes down on the Lawful side of things, and after Leo murders the king, Diego prepares to avenge him.
• Uncertain Doom: Leo fights him, and when Verm comes upon the scene soon after Diego's hammer is lying abandoned, with No Body Left Behind. In the epilogue the other, more normal guard who was posted next to Diego at the entrance to the throne room is sad and doesn't want to talk about him, so he's probably dead.
• The Unfought: Despite his unique design and position all screaming 'boss fight', his only fight is with Leo, who defeats him offscreen. The OST includes unused music for this battle.

Plaguemaster Aquila

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• Despair Event Horizon: Aquila crossed this long before the game's events, when they read a book that detailed the irreparable destruction wrought on the environment by humankind. It was this that convinced them that the both the human and rodent worlds were beyond salvation.
• Flunky Boss: Their boss fight has them summon Mecha-Mooks to aid them in battle and help stall the party.
• For Science!: The stated reason that they don't seem to present as any particular gender is that they consider gender to be a distraction from what really matters: Science.
• Grumpy Old Man: Aquila is well aware of how little time they have left and resent how the Gods, being so Long-Lived, are afforded enough time to get to make mistakes. They also despise most other rodents, having scornful things to say about all strata of society.
• Mad Scientist: Subverted. At first all mentions of them paint them as an extremely sinister figure, with occasional mentions made of sinister experiments (including one that got them kicked out of the royal court), but the experiments that they have Siobhan help with are pretty tame - making mouse robots. Aquila attributes their bad reputation to anti-intellectualism. They may have pivoted away from experimenting on rodents by then. While they do end up being an antagonist, it's for reasons largely unrelated to their scientific endeavors.
• Noodle Incident: What exactly got them thrown out of the royal court in the first place.
• Power Floats: When the party fights them, Aquila casts off their cloak and takes a Standard Power Up Pose, suspended off the ground on their tail.
• Short-Lived Organism: All rodents are compared to the Gods, but Aquila is intensely, resentfully aware of it in a way the other characters aren't, frequently bemoaning that a rodent's life is short and meaningless. This is perhaps attributable to them reading human-made books and internalizing some of the way humans talk about time.
• Time-Limit Boss: The battle against Plaguemaster Aquila has a four-minute time limit before they launch Excalibur, killing a huge portion of Rodentia.

Titans

Tiger

A housecat who's been terrorizing Cranbaile.

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• Advertised Extra: The game's main promotional art features him, but he's a Disc-One Final Boss who you can optionally return to and re-fight later in the game.
• Panthera Awesome: He's just a normal house cat, but in proportion to the rodent protagonists Tiger is absolutely colossal.

The Kraken

An octopus that has taken up residence in the sewers.

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• Combat Tentacles: It can lash at people with its tentacles.
• Kraken and Leviathan: While it's not a very large octopus it's quite big compared to the rodent cast and calls itself the Kraken.
• Rhymes on a Dime: For some reason, it speaks exclusively in rhyming verse.

The Curse of Solhill

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• Dark Is Evil: It's fought shrouded in darkness in a pitch-black room.
• The Darkness Gazes Back: The first sight of it are its green eyes.
• Maybe Magic, Maybe Mundane: Rosalie and Maisie seem to see it as a genuinely supernatural beast, but Verm think's it's probably just an escaped human pet. The setting does have some supernatural elements, but aside from London being an extremely odd place to find a hooded cobra it doesn't seem to do anything really unusual for a snake. It is unique among the Titans in that it doesn't speak - the Wizard Lizard, a legless lizard resembling a snake, can, so you'd think snakes could as well.
• Optional Boss: It's only fought if you choose to go to Solhill near the end of the game.
• Snakes Are Sinister: It's a cobra twined about the skull of a fox, barely visible in the darkness.

The Cailleach

An ancient seer who lives at the top of Ashen Tower, near Sky Garden.

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• Dead All Along: Murdered by Lamia seasons ago at the Duke's order, after she prophesied he would be overthrown. She's survived by four fledglings, which retain a portion of her knowledge.
• Long-Lived: Said to be four hundred seasons old, a hundred years in human terms and well beyond the lifespan of a normal owl.
• The Owl-Knowing One: She's an owl Titan said to be omniscient and to have the gift of prophecy. Verm seeks her out for more information on the Yellow God.
• The Reveal: That she was an owl doesn't come up for a while.

Nightwings

The Cailleach's four children, Bane-Ghial, Sheer-Ghlass, Doo-Dorraghey, and Jiarg-Lossanagh, who have taken her place atop Ashen Tower.

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• Horsemen of the Apocalypse: For some reason, their colors (and names) match the horses of the biblical Horsemen. Bane-Ghial is white (Conquest), Sheer-Ghlass is green (Death), Doo-Dorraghey is black (Famine), and Jiarg-Lossanagh is red (War). The party rides them into battle in the opening of their attack on Sky Garden.
• Ominous Owl: The Cailleach herself doesn't seem to have been a fearful figure, or at least, other characters talk about her as being wise and ancient and don't mention that she was an owl, and there was no mention about her expecting tribute after answering questions. Her children are introduced as glowing eyes in a pitch black room and want to eat the party.
• The Owl-Knowing One: Like their mother before them, they claim to have some kind of supernatural knowledge.
• Pet the Dog: The player can have Verm's one question be about the Yellow God and how to kill it, or about his brother Lance and whether Lance forgives him. The Nightwing who answers in the latter case is fairly sympathetic about it.
• The Psycho Rangers: They mirror the composition of Verm's party: Bane-Ghial is their main melee fighter, Jiarg-Lossanagh attacks from range like Siobhan, Sheer-Ghlass inflicts status effects similarly to Gwen, and Doo-Dorraghey sings the same songs as Bruce.

The Gods

In General

• Giant Hands of Doom: Due to the massive size difference, the two battles against gods are really just against the gods' hands.
• Humans Are Cthulhu: The people of Rodentia call humanity gods, and many are terrified of even the chance of gaining their attention. The opening lets us briefly control a mouse who is attempting to attack one of them as a distraction, and it's an utter Curb-Stomp Battle where the human is left completely unharmed and the mouse is crushed to death almost immediately by an attack that does hundreds of times more damage than their maximum health.

The White God

• Ambiguous Gender: The party never sees its face and doesn't gender it in any way. The Yellow God, in comparison, is sometimes referred to as Him.
• Animal Testing: This is one of several white-coated scientists who's been performing tests on Lab Pets like Gwen and George.
• Background Boss: Before battle commences you can see it approaching in the background, pulling on white gloves, though when it's closer all you can see of it is its hands fading into darkness.
• Improbable Weapon User: It wields a scalpel in one of its hands.
• Karma Houdini: Verm insists they fight it to give Gwen "closure", as she's been harmed by the White Gods before escaping the lab. The party can defeat it, meaning that it stops trying to grab and slash them and walks away - walks, not runs. Verm, briefly despairing, says they barely scratched it.
• Optional Boss: Only fought if the player decides to return to Whitehall in the endgame.
• Superboss: The White God is one of the most resilient and by far the strongest opponent in the game, with attacks capable of causing a Total Party Kill if they go in unprepared.

The Yellow God

• Book Ends: The first encounter with the Yellow God takes place in an area introduced with the text "Heaven: Tread Softly." The Final Boss takes place in an area introduced with "Hell: Tread Softly."
• Hazmat Suit: The Yellow God of Death is, in actuality, just an ordinary human exterminator in a protective yellow suit so he isn't exposed to the Deadly Gas he uses for his job.
• Hopeless Boss Fight: Lance's battle with it in the prologue. None of his attacks deal a single point of damage to it, and once Lance gets its attention, the Yellow God uses its hand to crush Lance several times for damage way over his maximum health, killing him.
• The Power of Friendship: The Yellow God as the Final Boss can grab a character and attempt to subject them to Lance's fate in the prologue by repeatedly crushing them... but this time round, you have allies and dealing enough damage (including bleed damage) to the hand that grabbed them will release the character, saving them. Also, while even together the party's attacks just scratch the Yellow God's hands without really threatening it, they do manage to collectively heave the ceremonial mace of the House of Commons at it and knock it over.
• Reflective Eyes: When it's lying on the ground Verm goes up to its faceplate and sees himself reflected in its eyes.
• Screw This, I'm Outta Here: After Verm and his party land the final blow against the Yellow God, Verm hops over to meet him face-to-face and menacingly tells him to run. The following cutscene makes it pretty obvious that the Yellow God took Verm's advice.
• Seemingly Hopeless Boss Fight: The final battle against the Yellow God has Verm's party attacking the boss' feet, which have ??? HP each and damaging them doesn't put a dent in their health bar. Verm fortunately decides that the strategy isn't working and they need to get higher, causing the party to fight the boss' much more vulnerable hands.

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